Edinburgh – Centre Wesley Fofana has been ruled out of France’s Rugby World Cup warm-up Test with Scotland this weekend and has been replaced by Sofiane Guitoune, the French Rugby Federation announced on Friday.

Vice-captain Fofana, 31, is suffering from a thigh problem and 30-year-old Guitoune is in line to make his first Test appearance since the 2015 tournament.

Guitoune scored 16 tries in 31 matches for Toulouse last season as they won a record 20th Top 14 title and reached the European Champions Cup semi-finals.

The last of his five international caps, which all came on the wing, was against Romania four years ago and he joins Gael Fickou in Les Bleus’ midfield for Saturday’s match at Murrayfield.

Teams:

Scotland

15 Stuart Hogg, 14 Tommy Seymour, 13 Chris Harris, 12 Pete Horne, 11 Sean Maitland, 10 Finn Russell, 9 Greig Laidlaw (captain), 8 Blade Thomson, 7 Hamish Watson, 6 Ryan Wilson, 5 Sam Skinner, 4 Scott Cummings, 3 Willem Nel, 2 George Turner, 1 Gordon Reid

Substitutes: 16 Grant Stewart, 17 Allan Dell, 18 Simon Berghan, 19 Grant Gilchrist, 20 John Barclay, 21 George Horne, 22 Rory Hutchinson, 23 Blair Kinghorn

France:

15 Thomas Ramos; 14 Damian Penaud, 13 Gael Fickou, 12 Sofiane Guitoune, 11 Alivereti Raka; 10 Camille Lopez, 9 Antoine Dupont; 8 Charles Ollivon, 7 Gregory Alldritt, 6 Arthur Iturria; 5 Sebastien Vahaamahina, 4 Felix Lambey; 3 Rabah Slimani, 2 Guilhem Guirado (capt), 1 Jefferson Poirot

Substitutes: 16 Camille Chat, 17 Cyril Baille, 18 Emerick Setiano, 19 Romain Taofifenua, 20 Yacouba Camara, 21 Baptiste Serin, 22 Romain Ntamack, 23 Maxime Medard